Rookie co-pilot Scott faces his worst nightmare when a faltering Curtiss C-46 Commando engine and low fog force a scary landing. Ambitious young flyer Alex must prove himself when Buffalo's chief pilot gives him control of the 20-ton Douglas DC-4 for the very first time. And the boss' son Mikey gets in hot water when Buffalo Joe needs the backup Douglas DC-3 - and it's nowhere near ready.

Christmas is just around the corner, but Buffalo's crucial food and mail runs come to a halt. An engine seizure strands Scott, AJ and the C-46 plane, creating a chaotic chain reaction. Crusty mechanic Chuck must attempt a bare-handed engine swap in minus 30, and pressure from customers and co-workers pushes cargo manager Kelly to the breaking point.

A string of bad luck turns chief pilot Arnie's 66th birthday run into the flight from hell - and threatens to spoil his naughty birthday surprise. Ramp hand Jeremy is desperate to move up the ranks, but a female newbie could throw a wrench in his plans. And after the crew works all night to turn the DC- 4 into a flying gas tanker, new co-pilot Alex flies a tricky fuel haul.

When a power failure leaves Rankin Inlet exposed to the brutal northern elements, captain Justin races to deliver an emergency generator. Ramp hand Jeremy tries to move ahead of co-worker Audrey by training a new hire from India to replace him. And when the recession nips at Joe's heels, he tries to broker a $7-million deal that could save Buffalo's hide.

Mechanic Cory races against the clock to turn two short-haul waterbombers into transatlantic aircraft for the epic journey to Turkey. It's up to engineer Adam to get the C-46 crew out of a jam on a remote airstrip in minus 40. And when captain Devan and co-pilot Scott encounter the menace of wind shear at notorious Bear Rock, it's a hair-raising experience that tests their friendship.

A charter flight falls through the cracks after the McBryans attend a wild family wedding, and an irate Joe pulls captain Justin from the Turkey test flights to do the job. Justin's task is to move an impatient Brit, his dog and all his worldly possessions to a remote northern outpost, but the weather isn't cooperating. Meanwhile, Devan ponders a job offer in Africa...

A furious Joe fights against a federal suspension that will prevent him from carrying passengers for the first time in 20 years. It's also a setback for Jeremy and the other rampies, which takes a turn for the worse with an unexpected injury. For the Turkey waterbomber team it's finally a go.

AJ and Candace get married in the Ice Castle. A frustrated Chuck tries to get the Electra to pay her way. A flummoxed Mikey tries to bake a cake for Joe's birthday. And in Newfoundland, a fuel leak, a sobering survival course and a tragic crash rattle the Turkey crew. When raging winds provide the long-awaited weather window, will they chance on the treacherous North Atlantic crossing?

The Turkey crew makes the critical jump over the North Atlantic. But en route to Portugal, one crew is forced to turn back. With no power on one engine and no way to maintain altitude, they must get back into Santa Maria, where the airport is on full emergency alert. And Mikey and Scott search for a new rampie, while Jeremy's hopes plummet even further.

C-46 and DC-4 crews race against the spring melt to fly mining equipment to an airstrip on a frozen lake. Jeremy tries to catch up to fellow rampies Audrey and Wilf. And on the final leg to Turkey, Arnie and Justin face some serious volcanoes and busy Mediterranean air traffic.

After thousands of hours of painstaking preparation and dangerous flying, a terrible accident devastates the waterbomber crew in Turkey - and everyone at Buffalo. Wilf's first run as flight attendant raises eyebrows among the staff. And a Scottish airplane buff fulfills the dream of a lifetime when Joe lets him ride in the captain's seat of his DC-3.

When her husband quits after a blowout with Joe, Kelly is put in a terrible position. Will she stay at Buffalo - or will she go? Scott gets the news he must stand down from flying so another co-pilot can fast-track to captain. He pulls out all the stops to secure a new charter, but his rival also wants to fly the new job. Jeremy has a chance to beat the other rampies to a job piloting the DC-3.

After her husband's acrimonious departure from Buffalo, Kelly has a confrontation that decides whether she, too, will quit. Jeremy makes a mistake that could end his Buffalo career. An aviation gas shortage brings a sobering dose of reality to the McBryans. And after a visit from his mom, fate throws Scott an unexpected curve ball. When the dust settles, someone will be leaving Buffalo Airways.